And now, after putting the stomach contents through a battery of tests, the researchers determined the ice mummy's final meal: dried ibex meat and fat, red deer, einkorn wheat, and traces of toxic fern.
By lining up the position of surrounding organs in radiographic images, the team finally found the stomach.
View Images A tiny piece of undigested plant tissue researchers spotted in the Iceman's stomach contents.
But curiously, though DNA analysis suggests red deer (Cervus elaphus) was also part of the meal, researchers couldn't figure out what part of the creature Ötzi ate.
Ötzi also ate einkorn wheat and the toxic bracken fern.